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Camacho, O., Fernández‐García, C., Vieira, C., Gurgel, C. F. D., Norris, J. N., Freshwater, D. W., & Fredericq, S. (2019). The systematics of Lobophora (Dictyotales, Phaeophyceae) in the western Atlantic and eastern Pacific oceans: eight new species. Journal of phycology. DOI: 10.1111/jpy.12850

Collin, R., Venera‐Pontón, D. E., Driskell, A. C., Macdonald, K. S., Chan, K. Y. K., & Boyle, M. J. (2019). Documenting neotropical diversity of phoronids with DNA barcoding of planktonic larvae. Invertebrate Biology, e12242. DOI: 10.1111/ivb.12242

Collin, R., Venera-Pontón, D., Driskell, A., Macdonald, K., & Boyle, M. (2019). Planktotrophic Brachiopod Larvae from the Pacific and Caribbean of Panama. Diversity11(1), 2. DOI: 10.3390/d11010002

Curini-Galletti, M., Stocchino, G. A., & Norenburg, J. L. (2019). New species of Duplominona Karling, 1966 and Pseudominona Karling, 1978 (Platyhelminthes: Proseriata) from the Caribbean. Zootaxa4657(1), zootaxa-4657. DOI: 10.11646/zootaxa.4657.1.5

Jaime G. Morín, E. Venera-Pontón Dagoberto, C. Driskell Amy, A. Sánchez Juan, R. Lasker Howard, Collin Rachel (2019). Reference DNA barcodes and other mitochondrial markers for identifying Caribbean Octocorals. Biodiversity Data Journal7, e30970. DOI: 10.3897/BDJ.7.e30970

Oliveira, L. M., Hoeksema, B. W., & Rocha, R. M. (2019). Polysyncraton (Ascidiacea, Didemnidae): a re-examination of some specimens and descriptions of three new species. European Journal of Taxonomy2019(519), 1-25. DOI: 10.5852/ejt.2019.519

Rocha, R. M., & Counts, B. K. (2019). Pyura (Tunicata: Ascidiacea: Pyuridae) on the coasts of Panama. Zootaxa4564(2), 491-513. DOI: 10.11646/zootaxa.4564.2.9

Skinner, L. F., Rocha, R. M., & Counts, B. K. (2019). Pyura gangelion and Pyura beta sp.nov. (Ascidiacea: Pyuridae): an exotic and a new tunicate from the West Atlantic. Zootaxa4545(2), 264-276. DOI: 10.11646/zootaxa.4545.2.6

Vicente, J., Ríos, J. A., Zea, S., & Toonen, R. J. (2019). Molecular and morphological congruence of three new cryptic Neopetrosia spp. in the Caribbean. PeerJ7, e6371. DOI: 10.7717/peerj.6371

Worsaae, K., Mikkelsen, M. D., & Martínez, A. (2019). Description of six new species of Mesonerilla (Nerillidae, Annelida) and an emended description of M. intermedia Wilke, 1953, from marine and cave environments. Marine Biodiversity, 1-25. DOI: 10.1007/s12526-019-00984-6



Anker, A. (2018). Alpheus naranjo, a new brightly coloured snapping shrimp from the Caribbean coast of Panama (Malacostraca, Decapoda, Alpheidae). Zootaxa4433(3), 574-582. DOI: 10.11646/zootaxa.4433.3.11

Medrano, S., Krug, P. J., Gosliner, T. M., Kumar, A. B., & Valdés, Á. (2018). Systematics of Polybranchia Pease, 1860 (Mollusca: Gastropoda: Sacoglossa) based on molecular and morphological data. Zoological Journal of the Linnean Society. DOI: 10.1093/zoolinnean/zly050

Miglietta, M. P., Maggioni, D., & Matsumoto, Y. (2018). Phylogenetics and species delimitation of two hydrozoa (phylum Cnidaria): Turritopsis (McCrady, 1857) and Pennaria (Goldfuss, 1820). Marine Biodiversity, 1-16. DOI: 10.1007/s12526-018-0891-8

Miglietta, M. P., Piraino, S., Pruski, S., Alpizar Gonzalez, M., Castellanos-Iglesias, S., Jerónimo-Aguilar, S., … & Moncada, A. (2018). An integrative identification guide to the Hydrozoa (Cnidaria) of Bocas del Toro, Panama. Neotropical Biodiversity4(1), 103-113. DOI: 10.1080/23766808.2018.1488656

Pachelle, P. P., Leray, M., Anker, A., & Lasley, Robert (2018). Five new records of marine shrimps (Decapoda: Caridea, Stenopodidea) from the Caribbean coast of Panama. Zootaxa4438(1), 128-136. DOI: 10.11646/zootaxa.4438.1.5

Paiva, S. V., & Rocha, R. M.(2018). A large marine Caribbean mushroom field: description of Eudistoma amanitum sp. nov.(Ascidiacea: Polycitoridae). Zootaxa4399(3), 443-450. DOI: 10.11646/zootaxa.4399.3.13



De Grave, S., & Anker, A. (2017). An annotated checklist of marine caridean and stenopodidean shrimps (Malacostraca: Decapoda) of the Caribbean coast of Panama. Nauplius25. DOI: 10.1590/2358-2936e2017015

Richards, J. L., Sauvage, T., Schmidt, W. E., Fredericq, S., Hughey, J. R., & Gabrielson, P. W. (2017). The coralline genera Sporolithon and Heydrichia (Sporolithales, Rhodophyta) clarified by sequencing type material of their generitypes and other species. Journal of phycology53(5), 1044-1059. DOI: 10.1111/jpy.12562

Valdés, Á., Breslau, E., Padula, V., Schrödl, M., Camacho, Y., Malaquias, M. A. E., … & Gosliner, T. M. (2017). Molecular and morphological systematics of Dolabrifera Gray, 1847 (Mollusca: Gastropoda: Heterobranchia: Aplysiomorpha). Zoological journal of the Linnean Society184(1), 31-65. DOI: 10.1093/zoolinnean/zlx099



Anker, A., & Caripe, J. V. (2016). Leptalpheus pereirai sp. nov., a new alpheid shrimp from Panama and Venezuela (Decapoda: Caridea). Zootaxa4127(1), 185-191. DOI: 10.11646/zootaxa.4127.1.11

Collin, R., Fredericq, S., Freshwater, D. W., Gilbert, E., Madrid, M., Maslakova, S., … & Thacker, R. W. (2016). TaxaGloss-A glossary and translation tool for biodiversity studies. Biodiversity Data Journal, (4). DOI: 10.3897/BDJ.4.e10732

Goodheart, J. A., Ellingson, R. A., Vital, X. G., Galvão Filho, H. C., McCarthy, J. B., Medrano, S. M., … & Hoover, C. A. (2016). Identification guide to the heterobranch sea slugs (Mollusca: Gastropoda) from Bocas del Toro, Panama. Marine Biodiversity Records9(1), 56. DOI: 10.1186/s41200-016-0048-z

Grajales, A., & Rodríguez, E. (2016). Elucidating the evolutionary relationships of the Aiptasiidae, a widespread cnidarian–dinoflagellate model system (Cnidaria: Anthozoa: Actiniaria: Metridioidea). Molecular phylogenetics and evolution94, 252-263. DOI: 10.1016/j.ympev.2015.09.004

Hernandez-Kantun, J. J., Gabrielson, P., Hughey, J. R., Pezzolesi, L., Rindi, F., Robinson, N. M., … & Adey, W. (2016). Reassessment of branched Lithophyllum spp. (Corallinales, Rhodophyta) in the Caribbean Sea with global implications. Phycologia55(6), 619-639. DOI: 10.2216/16-7.1

Iha, C., O’Shaughnessy, K. A., Guimarães, S. M., Oliveira, M. C., & Freshwater, D. W. (2016). Taxonomic reappraisal of Gelidium coarctatum (Gelidiales, Rhodophyta) and Gelidium lineare sp. nov. from the tropical western Atlantic. Phycologia55(5), 555-563. DOI: 10.2216/16-17.1

Krug, P. J., Vendetti, J. E., & Valdés, Á. (2016). Molecular and morphological systematics of Elysia Risso, 1818 (Heterobranchia: Sacoglossa) from the Caribbean region. Zootaxa4148(1), 1-137. DOI: 10.11646/zootaxa.4148.1.1

Pardos, F., Herranz, M., & Sánchez, N. (2016). Two sides of a coin: the phylum Kinorhyncha in Panama. II) Pacific Panama. Zoologischer Anzeiger-A Journal of Comparative Zoology265, 26-47. DOI: 10.1016/j.jcz.2016.06.006

Pardos, F., Sánchez, N., & Herranz, M. (2016). Two sides of a coin: the phylum Kinorhyncha in Panama. I) Caribbean Panama. Zoologischer Anzeiger-A Journal of Comparative Zoology265, 3-25. DOI: 10.1016/j.jcz.2016.06.005

Valdez, A., Medrano, S., & Bhave, V. (2016). A new species of Cuthona Alder and Hancock, 1855 (Gastropoda: Heterobranchia: Nudibranchia: Tergipedidae) from the Caribbean Sea. Nautilus130(2), 72-78.



Easson, C. G., Matterson, K. O., Freeman, C. J., Archer, S. K., & Thacker, R. W. (2015). Variation in species diversity and functional traits of sponge communities near human populations in Bocas del Toro, Panama. PeerJ3, e1385. DOI: 10.7717/peerj.1385

Martínez, A., Di Domenico, M., Rouse, G. W., & Worsaae, K. (2015). Phylogeny and systematics of Protodrilidae (Annelida) inferred with total evidence analyses. Cladistics31(3), 250-276. DOI: 10.1111/cla.12089



Di Domenico, M., Martínez, A., Lana, P., & Worsaae, K. (2014). Molecular and morphological phylogeny of Saccocirridae (Annelida) reveals two cosmopolitan clades with specific habitat preferences. Molecular Phylogenetics and Evolution75, 202-218. DOI: 10.1016/j.ympev.2014.02.003

Kantun, J. J. H., Riosmena-Rodriguez, R., Adey, W. H., & Rindi, F. (2014). Analysis of the cox 2-3 spacer region for population diversity and taxonomic implications in rhodolith-forming species (Rhodophyta: Corallinales). Phytotaxa190(1), 331-354. DOI: 10.11646/phytotaxa.190.1.20

Leasi, F., & Norenburg, J. L. (2014). The necessity of DNA taxonomy to reveal cryptic diversity and spatial distribution of meiofauna, with a focus on Nemertea. PLoS one9(8), e104385. DOI: 10.1371/journal.pone.0104385

Olson, J. B., Thacker, R. W., & Gochfeld, D. J. (2014). Molecular community profiling reveals impacts of time, space, and disease status on the bacterial community associated with the Caribbean sponge Aplysina cauliformisFEMS microbiology ecology87(1), 268-279. DOI: 10.1111/1574-6941.12222

Redmond, N. E., Morrow, C. C., Thacker, R. W., Diaz, M. C., Boury-Esnault, N., Cárdenas, P., … & Kayal, E. (2013). Phylogeny and systematics of Demospongiae in light of new small-subunit ribosomal DNA (18S) sequences. Integrative and comparative biology53(3), 388-415. DOI: 10.1093/icb/ict078

Sal-Moyano, M., Lagos-Tobias, A., Felder, D. L., & Mantelatto, F. L. (2014). Relative growth and reproductive parameters in a population of Microphrys bicornutus (Brachyura, Majoidea) from Bocas del Toro, Caribbean Sea, Panama. Revista de Biología Marina y Oceanografía. DOI:

Thoma, B. P., Guinot, D., & Felder, D. L. (2014). Evolutionary relationships among American mud crabs (Crustacea: Decapoda: Brachyura: Xanthoidea) inferred from nuclear and mitochondrial markers, with comments on adult morphology. Zoological Journal of the Linnean Society170(1), 86-109. DOI: 10.1111/zoj12093



Alvizu, A., Díaz, M. C., Bastidas, C., Rützler, K., Thacker, R. W., & Márquez, L. M. (2013). A skeleton‐less sponge of Caribbean mangroves: invasive or undescribed?. Invertebrate Biology132(2), 81-94. DOI: 10.1111/ivb.12015

Cañete, J. I., & Rocha, R. M. (2013). Modiolarca lateralis (Pteryomorphia: Mytilidae): Bivalvo asociado a seis especies de ascidias de Bocas del Toro, Panamá. Latin american journal of aquatic research41(5), 1030-1035. DOI: 103856/vol41-issue5-fulltext-25

Diaz, M. C., Thacker, R. W., Redmond, N. E., Matterson, K. O., & Collins, A. G. (2013). Phylogenetic novelties and geographic anomalies among tropical VerongidaIntegrative and comparative biology53(3), 482-494. DOI: 10.1093/icb/ict033

Freeman, C. J., Thacker, R. W., Baker, D. M., & Fogel, M. L. (2013). Quality or quantity: is nutrient transfer driven more by symbiont identity and productivity than by symbiont abundance?. The ISME journal7(6), 1116. DOI: 10.1038/ismej.2013.7

Hajdu, E., de Paula, T. S., Redmond, N. E., Cosme, B., Collins, A. G., & Lôbo-Hajdu, G. (2013). Mycalina: another crack in the Poecilosclerida framework. Integrative and Comparative Biology53(3), 462-472. DOI: 10.1093/icb/ict074

Hirose, M., & Hirose, E. (2013). Photosymbiotic ascidians from oceanic islands in the tropical Pacific as candidates of long-dispersal species: morphological and genetic identification of the species. Aquatic Invasions8(3). DOI: 10.3391/ai.2013.8.3.03

Martínez, A., Di Domenico, M., Jörger, K., Norenburg, J., & Worsaae, K. (2013). Description of three new species of Protodrilus (Annelida, Protodrilidae) from Central America. Marine Biology Research9(7), 676-691. DOI: 10.1080/17451000.2013.765574

Page, T. J., Torati, L. S., Cook, B. D., Binderup, A., Pringle, C. M., Reuschel, S., … & Hughes, J. M. (2013). Invertébrés sans frontières: large scales of connectivity of selected freshwater species among Caribbean Islands. Biotropica45(2), 236-244. DOI: 10.1111/j.1744-7429.2012.00900.x

Pérez-Portela, R., Arranz, V., Rius, M., & Turon, X. (2013). Cryptic speciation or global spread? The case of a cosmopolitan marine invertebrate with limited dispersal capabilities. Scientific Reports3, 3197. DOI: 10.1038/srep03197

Thacker, R. W., Hill, A. L., Hill, M. S., Redmond, N. E., Collins, A. G., Morrow, C. C., … & Hall, C. (2013). Nearly complete 28S rRNA gene sequences confirm new hypotheses of sponge evolution. Integrative and Comparative Biology53(3), 373-387. DOI: 10.1093/icb/ict071

Wirshing, H. H., Feldheim, K. A., & Baker, A. C. (2013). Vectored dispersal of Symbiodinium by larvae of a Caribbean gorgonian octocoral. Molecular ecology22(17), 4413-4432. DOI: 10.1111/mec.12405



Brown, F. D., & Swalla, B. J. (2012). Evolution and development of budding by stem cells: ascidian coloniality as a case study. Developmental biology369(2), 151-162. DOI: 10.1016/j.ydbio.2012.05.038

Hirose, E., Turon, X., López-Legentil, S., Erwin, P. M., & Hirose, M. (2012). First records of didemnid ascidians harbouring Prochloron from Caribbean Panama: Genetic relationships between Caribbean and Pacific photosymbionts and host ascidians. Systematics and Biodiversity10(4), 435-445. DOI: 10.1080/14772000.2012.735716

Mamoozadeh, N. R., & Freshwater, D. W. (2012). Polysiphonia sensu lato (Ceramiales, Florideophyceae) species of Caribbean Panama including Polysiphonia lobophoralis sp. nov. and Polysiphonia nuda sp. nov. Botanica marina55(4), 317-347. DOI: 10.1515/bot-2012-0147

Torati, L. S., & Mantelatto, F. L. (2012). Ontogenetic and evolutionary change of external morphology of the neotropical shrimp Potimirim (Holthuis, 1954) explained by a molecular phylogeny of the genus. Journal of Crustacean Biology32(4), 625-640. DOI: 10.1163/193724012X635322



Bonnet, N. Y., & Rocha, R. M. (2011). The family Ascidiidae Herdman (Tunicata: Ascidiacea) in Bocas del Toro, Panama. Description of six new species. Zootaxa2864, 1-33. DOI: 10.11646/zootaxa.2864.1.1

Freeman, C. J., & Thacker, R. W. (2011). Complex interactions between marine sponges and their symbiotic microbial communities. Limnology and Oceanography56(5), 1577-1586. DOI: 10.4319/lo.2011.56.5.1577

Hochberg, R., & Atherton, S. (2011). A new species of Lepidodasys (Gastrotricha, Macrodasyida) from Panama with a description of its peptidergic nervous system using CLSM, anti-FMRFamide and anti-SCPB. Zoologischer Anzeiger-A Journal of Comparative Zoology250(2), 111-122. DOI: 10.1016/j.jcz.2010.12.002

Krayesky, D. M., Norris, J. N., West, J. A., & Fredericq, S. (2011). The Caloglossa leprieurii complex (Delesseriaceae, Rhodophyta) in the Americas: the elucidation of overlooked species based on molecular and morphological evidence. Cryptogamie, Algologie32(1), 37-63. DOI: 10.7872/crya.v32.iss1.2011.037

Shenkar, N., & Swalla, B. J. (2011). Global diversity of Ascidiacea. PLoS One6(6), e20657. DOI: 10.1371/journal.pone.0020657

Torati, L. S., Page, T., De Grave, S., & Anker, A. (2011). Atyidae and Palaemonidae (Crustacea: Decapoda: Caridea) of Bocas del Toro, Panama. Check List7, 798. DOI: 10.15560/11026



Collin, R., Mobley, A. S., Lopez, L. B., Leys, S. P., Diaz, M. C., & Thacker, R. W. (2010). Phototactic responses of larvae from the marine sponges Neopetrosia proxima and Xestospongia bocatorensis (Haplosclerida: Petrosiidae). Invertebrate Biology129(2), 121-128. DOI: 10.1111/j.1744-7410.2010.00196.x

Hochberg, R., & Atherton, S. (2010). Acanthodasys caribbeanensis sp. n., a new species of Thaumastodermatidae (Gastrotricha, Macrodasyida) from Belize and Panama. Zookeys61(1). DOI: 10.3897/zookeys.61.552



Cardenas, P., Menegola, C., Rapp, H. T., & Diaz, M. C. (2009). Morphological description and DNA barcodes of shallow-water Tetractinellida (Porifera: Demospongiae) from Bocas del Toro, Panama, with description of a new species. Zootaxa2276, 1-39. DOI:

Cárdenas, P., Rapp, H. T., Schander, C., & Tendal, O. S. (2010). Molecular taxonomy and phylogeny of the Geodiidae (Porifera, Demospongiae, Astrophorida) combining phylogenetic and Linnaean classification. Zoologica scripta39(1), 89-106. DOI: 10.1111/j.1463-6409.2009.00402.x

De Grave, S. (2009). A further sponge-dwelling species of the Periclimenes iridescens complex from the western Atlantic (Decapoda, Caridea, Palaemonidae). Crustaceana, 829-836. DOI:

Sánchez, J. A. (2009). Systematics of the candelabrum gorgonian corals (Eunicea Lamouroux; Plexauridae; Octocorallia; Cnidaria). Zoological Journal of the Linnean Society157(2), 237-263. DOI: 10.1111/j.1096-3642.2008.00515.x

Sperling, E. A., Peterson, K. J., & Pisani, D. (2009). Phylogenetic-signal dissection of nuclear housekeeping genes supports the paraphyly of sponges and the monophyly of Eumetazoa. Molecular biology and evolution26(10), 2261-2274. DOI: 10.1093/molbev/msp148



Erwin, P. M., & Thacker, R. W. (2008). Cryptic diversity of the symbiotic cyanobacterium Synechococcus spongiarum among sponge hosts. Molecular Ecology17(12), 2937-2947. DOI: 10.1111/j.1365-294X.2008.03808.x

Erwin, P. M., & Thacker, R. W. (2008). Phototrophic nutrition and symbiont diversity of two Caribbean sponge–cyanobacteria symbioses. Marine Ecology Progress Series362, 139-147. DOI: 10.3354/meps07464



Diaz, M. C., Thacker, R. W., Rützler, K., & Piantoni, C. (2007). Haliclona (Soestella) walentinae (Chalinidae, Haplosclerida) and Xestospongia bocatorensis (Petrosiidae, Haplosclerida), two new sponge species from Caribbean Panamá with filamentous cyanobacterial symbionts. Porifera Research-Biodiversity, Innovation and Sustainability, 31-39.

Erwin, P. M. & Thacker, R. W.. (2007). Incidence and identity of photosynthetic symbionts in Caribbean coral reef sponge assemblages. Journal of the Marine Biological Association of the United Kingdom, 87(6), 1683-1692. DOI: 10.1017/S0025315407058213

Erwin, P. M., & Thacker, R. W. (2007). Incidence and identity of photosynthetic symbionts in Caribbean coral reef sponge assemblages. Journal of the Marine Biological Association of the United Kingdom87(6), 1683-1692. DOI: 10.1017/S0025315407058213

Erwin, P. M., & Thacker, R. W. (2007). Phylogenetic analyses of marine sponges within the order Verongida: a comparison of morphological and molecular data. Invertebrate Biology126(3), 220-234. DOI: 10.1111/j.1744-7410.2007.00092.x

Gochfeld, D.J., C. Schlöder, & Thacker, R.W. (2007). Sponge community structure and disease prevalence on coral reefs in Bocas del Toro. Porifera Research: Biodiversity, Innovation, and Sustainability, 335-343.

Thacker, R. W., Diaz, M. C., Rützler, K., Erwin, P. M., Kimble, S. J., Pierce, M. J., & Dillard, S. L. (2007). Phylogenetic relationships among the filamentous cyanobacterial symbionts of Caribbean sponges and a comparison of photosynthetic production between sponges hosting filamentous and unicellular cyanobacteria. Porifera research: Biodiversity, innovation and sustainability, 621-626.